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March 21st, 2005
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Q. I read about the in pool insect drift occurring about 1 hour after dark and again early morning before first light. Both times the insects were reported to crawl back to their niche areas. I would like to get your theory about the purpose of the drift since the material I read indicated that the drift was still under consideration as to purpose and effect. The reference material is located at "https://www.cffcm.org/streameco/streameco-sec2-drifting.htm" I have had some luck with the Ku fish since we talked. I also located the Kashe Stream (nicknamed the Crystal Palace of the Chuoshui River) that contains rainbows. Best regards, Mike aka: Rollcaster
A.
Hi Mike. I'm going to do several things in response to your
submission, including some rebuttal to the article
on pool drift that you cite above.
First, I'd like to refer you to an earlier
column that I did on drifting insects. If
you'll go to my recent column on FAOL, scroll
down to "Previous Stream Doctor Columns" and
then to the list under 2nd Quarter 2003, you'll
find a column titled "Drifting Organisms?"
If you'll open this, you'll find a column
I wrote for Trout magazine on drift
plus some other information on the subject.
It pretty well sums up what we know about
the three types of drift: active, behavioral,
and catastrophic.
Second, I want to address your specific
question on pool drift and the purpose
and effect of drift. Then, in case others
open the website you cite, I'd like to say
a few words on the information (misinformation?)
given in the article.
Drift can occur in pools, but it is mostly
related to riffles and faster moving water.
In fact, studies have shown a decrease in
drifting insects as water enters a pool and
the insects settle to the bottom. As to
the purpose and effect of drift, at least
the behavioral aspect results in a downstream
dispersal of organisms. Competition for space
and food resources can result in insects
actively moving off of a site or being forced
into the drift by aggressive competition from
other insects. The effect results in dispersal
of the insects throughout the stream, but as I
mention in the FAOL column, a "colonization
cycle" exists so that all insects don't end
up in the ocean; mature egg-laying females
fly upstream to deposit their eggs, thus
resulting in an existing population in the
headwaters. Part of the behavioral aspect of
drift occurs as emergers leave the bottom to
reach the surface; they can't help but being
carried in the current as drifters at this time.
Now to the article on the website. If you
read the column on drift on FAOL cited above,
you'll find that drift can be active, that is,
from insects actively leaving the bottom, but
this is not for the purpose of obtaining food from
the water column. It occurs when they leave
the bottom, for one reason or another, and are
swept up in the current; they aren't out for a
swim! Of the various methods used by aquatic
insects to feed (known as functional feeding
groups; see my FAOL column, 2nd quarter 2004,
"What do insects eat?"), none actively obtain
food particles by picking them out of the water
column. Thus, they do not feed on plankton
(free floating organisms), as inferred on the
website.
Knowledge on drifting insects is not relatively
new. Published studies on this phenomenon have
been available since 1972, and observations on
drift of insects related to fish feeding were
done as far back as the 1940s; a great deal
is known about the subject. The website
article also infers that drift only takes place
twice a night and then for only minutes at a
time; this is incorrect. Drift numbers in a
stream are positively correlated with darkness;
it increases and peaks soon after the sky darkens,
remains relatively high during the night, and
decreases again at dawn, with often a lesser
peak just before dawn.
In fact, one researcher even found that during
a cloudy night, drift numbers actually decreased
momentarily for a short period when the moon appeared
through the clouds for a brief period. There is
some periodicity in certain drifting organisms,
and I'm not disputing the fact that some particular
organism exhibits the pattern the author of the
website article described; what I'm saying is that
the general pattern is one of high drift numbers
during darkness and lower numbers during the day.
Note that I'm not saying that there is no drift
during the day; there is, or we'd be wasting a
lot of time fishing drifting nymph patterns!
To further emphasize why drifting numbers are
highest during darkness, observation by infrared
light of stone surfaces during darkness revealed
much higher numbers of insects on the upper
surfaces during darkness than during the day.
The obvious conclusion is that (1) the insects
feel less susceptibility to predators during
darkness, and (2) being on the upper surfaces
of the stones, they are more likely to be swept
into the water column; hence, higher numbers
during darkness.
There's a good general discussion of the drift
phenomenon in Tom Water's book Wildstream.
He was one of the early scientists who studied drift.
If you have a question, please feel free to contact me.
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