I would guess that the fish were starting to get a little more active since the bugs were getting more active. In the middle of the winter there isnt a lot of bug activity so even though the water temps were low the fish were feeding. I see the same thing on the Lower Madison which is a bit warmer than the temps you had. If there are bugs around the fish tend to be active on the surface. And I love fishing on those days where the air temp in is in the 30's and it is raining/snowing . The fish in the lower Madison where I fish will very rarely rise if the sun is on the water but when it is cloudy and or the water is shaded then they will feed on or just under the surface.
Just my two cents and I may be wrong.
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Take care everyone and cya around. Mark
Take care and cya around,
Mark